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A great white pelican, Pelecanus onocrotalus, on a tree top. Khwai Concession, Okavango Delta, Botswana
Two white rhinoceroses, Ceratotherium simum, walking in the dust at sunset. Kalahari, Botswana
A male lion, Panthera leo, patrolling. Khwai Concession, Okavango Delta, Botswana
A malachite kingfisher, Alcedo cristata. Moremi Game Reserve, Okavango Delta, Botswana
A sub-adult lion, Panthera leo, drinks at waterhole. Savuti, Chobe National Park, Botswana
Two white rhinoceroses, Ceratotherium simum, walking in a cloud of dust at sunset. Kalahari, Botswana
Portrait of a leopard, Panthera pardus, resting on a tree and looking at the camera. Khwai Concession, Okavango Delta, Botswana
Close-up portrait of a male lion, Panthera leo. Moremi Game Reserve, Okavango Delta, Botswana
A saddle-billed stork, Ephippiorhynchus senegalensis, walking. Moremi Game Reserve, Okavango Delta, Botswana
Three chacma baboons, Papio ursinus, sitting in a tree in Chobe National Park. Botswana
A chacma baboon, Papio ursinus, with a newborn in Chobe National Park. Botswana
Two African elephants, Loxodonta Africana, sparring in Okavango Delta's Khwai Concession. Botswana
An aerial view of a herd of African buffalo, Syncerus caffer. Okavango Delta, Botswana
African elephant, Loxodonta Africana, drinking in Okavango Delta's Khwai Concession. Botswana
An eastern yellow-billed hornbill, Tockus flavirostris, perching in the Okavango Delta's Khwai Concession. Botswana
A female leopard, Panthera pardus, walking in Chobe National Park's Savuti marsh. Botswana
A leopard, Panthera pardus, standing on a dead fallen tree in the Okavango Delta's Khwai Concession. Botswana
A lilac-breasted roller, Coracias caudata, perches on a tree branch. Khwai Concession, Okavango Delta, Botswana
An African elephant, Loxodonta Africana, drinking in Okavango Delta's Khwai Concession. Botswana
A yellow-billed stork, Mycteria ibis, in Chobe National Park. Botswana
A hippopotamus, Hippopotamus amphibius, in water, exhibiting territorial behavior. Khwai Concession Area, Okavango, Botswana
An African elephant, Loxodonta Africana, glancing at the photographer as it walks by. Chobe National Park, Botswana
Close-up portrait of a plains or Burchell's zebra, Equus burchellii. Khwai Concession Area, Okavango, Botswana
A male and a female southern giraffe, Giraffa camelopardalis, walking together through a shrubby landscape. Chobe National Park, Botswana
Close-up of a hippopotamus, Hippopotamus amphibius, grazing on a grass island. Chobe River, Chobe National Park, Botswana
A herd of African elephants, Loxodonta Africana, drinking. Mashatu Game Reserve, Botswana
A southern giraffe, Giraffa camelopardalis, with a one-week-old calf. Mashatu Game Reserve, Botswana
Vehicular tracks on a sandy dirt road through a vast savanna. Kudiakam Pan, Nxai Pan National Park, Botswana
Portrait of a southern giraffe, Giraffa camelopardalis, looking at the camera. Mashatu Game Reserve, Botswana
Portrait of a leopard, Panthera pardus, yawning. Mashatu Game Reserve, Botswana
Close-up portrait of a lion, Panthera leo. Mashatu Game Reserve, Botswana
Portrait of an alert hippopotamus, Hippopotamus amphibius, in the water. Khwai Concession Area, Okavango, Botswana
Portrait of a waterbuck, Kobus ellipsiprymnus. Khwai Concession, Okavango Delta, Botswana
Portrait of a pair of resting lions, Panthera leo. Mashatu Game Reserve, Botswana
Portrait of a giant kingfisher, Megaceryle maximus, perching in a bush. Chobe National Park, Botswana
A female African elephant, Loxodonta Africana, walking with her calf. Chobe National Park, Botswana
Portrait of a female southern giraffe, Giraffa camelopardalis, among shrubby trees. Savuti Marsh, Chobe National Park, Botswana
Close-up of an African elephant's feet, Loxodonta Africana. Mashatu Game Reserve, Botswana
Portrait of an African buffalo, Syncerus caffer, looking at the camera. Chobe National Park, Botswana
An African elephant calf, Loxodonta Africana, protected by its mother. Mashatu Game Reserve, Botswana
A female leopard, Panthera pardus, on a large tree branch looking at the camera. Khwai Concession, Okavango Delta, Botswana
Close-up of a hippopotamus, Hippopotamus amphibius, grazing on a grass. Chobe National Park, Botswana
A back-lit portrait of a chacma baboon, Papio ursinus, looking at the camera as it walks by. Chobe National Park, Botswana
An aerial view of plains zebras, Equus quagga, walking in a flood plain. Okavango Delta, Botswana
Portrait of a chacma baboon, Papio ursinus, looking at the camera as it walks by. Chobe National Park, Botswana